See the images

Biodiversity & Conservation

Use of Images

By contributing images (and/or video) to MarLIN, the owners of those images have agreed to their use on the MarLIN website, in MarLIN publications, posters and other promotional material. Contributors retain the copyright of their images. MarLIN indicates the individual or organization that supplied each image and brief contact details on the website.

Unless the image providers has stated to the contrary (see acknowledgement)

Images may be used:

Provided that: that users acknowledge the photographer, copyright holder and that it was published on the MarLIN website. For example

Image: Jon Davies/JNCC (published on the MarLIN website)

Images MUST NOT be used:

If images are required for commercial purposes or for wide dissemination, the image provider MUST be contacted for permission directly (unless otherwise stated and not through the MarLIN team) and the form of acknowledgement obtained from the owner. Contact details for image providers are provided on the MarLIN website as part of our acknowledgement.

Please direct any questions regarding use of the MarLIN images (inc. video) to Dr Harvey Tyler-Walters ( or care of MarLIN via

The MarLIN team would like to thanks the following photographers, scientists and organizations for providing images for the MarLIN Website.

Jim Anderson

Jim Anderson has been diving for 15 years with the Scotish Sub Aqua Club and has an exensive collection of Scottish underwater images, particularly invertebrates. For example visit the Scottish Nudibranchs Website.

Jim Anderson
Baird Hall
High Street, Linlithgow
West Lothian EH49 7ES

Lin Baldock

Lin Baldock is interested in the distribution of temperate marine invertebrates and has a collection of underwater images from the UK and Ireland.


MarLIN in the first instance

Guy Baker

Guy Baker works for the Marine Life Information Network as Communications and Outreach Officer. He is an amateur naturalist and takes pictures of marine life in his spare time.

Guy Baker

Peter Bardsley

Peter Bardsley is an amateur photographer, diver and conservationist with a keen interest and love of the underwater world which started as a teenager over 40 years ago.

Peter Bardsley

Peter Barfield

Peter Barfield is a marine biologist and freelance environmental consultant with a growing collection of digital images of both infaunal macrobenthos from UK samples and intertidal fauna from Cornwall and elsewhere.


Peter Barfield,
Sea-nature Studies
Tel: +44 (0)1208 851040

R.S.K. Barnes

Images reproduced with the author's permission from:
Barnes, R.S.K., 1994. The brackish-water fauna of northwestern Europe: An identification guide to brackish-water habitats, ecology and macrofauna for field workers, naturalists and students. Cambridge: Cambridge University Press.

Dr R.S.K.Barnes
Department of Zoology
University of Cambridge
Downing Street
Tel: +44 (0) 1223 333 296

Dan Bayley

Dan Bayley is a marine ecologist and works for the Marine Life Information Network and UK MPA centre at the MBA. In his spare time he is also a keen diver and photographer.


Robert Beharie

Robert Beharie works on shoreline wave action measurements and the determination environmental and ecological impacts from wave energy converter arrays. Robert continues monitoring shoreline wave action and carries out habitat and biotope surveys on the West coast of Orkney.

International Centre for Island Technology
Heriot Watt University
IPE Orkney Campus
Academy Buildings
Back Road
Orkney, KW16 3AW
Tel: +44 (0)1856 850605
Fax: +44 (0)1856 851349

Dr James Bell

Dr James Bell is a senior lecturer in marine biology at the Victoria University of Wellington, New Zealand whose research interests focus on Marine Protected Areas, self-recruitment and rocky shore and sponge ecology.

Dr James Bell,
School of Biological Sciences,
PO Box 600,
Wellington 6140,
New Zealand.

Steve Bell

Steve Bell is a Mountaineer, Sea Kayaker and Sea Kayak Coach, Amateur photographer and occasional freelance writer.

Steve Bell
19 Station Gate,

Dr Matt Bentley

Dr Matt Bentley is a research scientist working at the University of Newcastle but does not normally offer picture agency agreements. MarLIN should be contacted if you wish to use commercially any of the images provided by Dr Bentley and included on the MarLIN Website.

Dr Matt Bentley,
Department of Marine Sciences and Coastal Management,
Ridley Building,
University of Newcastle,
Newcastle upon Tyne, NE1 7RU.

Dr Brian Bett

Dr Brian Bett is a marine biologist at the Southampton Oceanography Centre (SOC). The DEEPSEAS Group at SOC hold a very large collection of deep-sea floor photographs and video and a major national collection of deep-sea invertebrate specimen material ("Discovery Collections") from the waters around the British Isles and world-wide.  Images provided by Dr Brian Bett must not be used in any capacity without prior permission from Dr Brian Bett.

Dr Brian Bett / Mrs Pamela Talbot,
Southampton Oceanography Centre,
Empress Dock,
Southampton SO14 3ZH, UK.
Tel. +44 (0)23 80596355
Fax. +44 (0)23 80596247


BIODIDAC aims to:

For information about image sources and permission to use BIODIDAC material please visit the BIODIDAC Website

Dr John Bishop

John Bishop teaches at the University of Plymouth; his research centres on ascidians and bryozoans.

John Bishop
Marine Biological Association
Citadel Hill Laboratory
Plymouth PL1 2PB
Tel: +44 (0)1752 633229

Jeff Blincow

Jeff Blincow is an amateur naturalist interested in all forms of wildlife.

Jeff Blincow

Dr Gerald Boalch

Dr Gerald Boalch is a marine botanist and Editor-in-Chief of the scientific journal 'Botanica Marina' ( He has a large collection of colour slides of seaweeds and phytoplankton, together with slides of the Torrey Canyon and Amoco Cadiz oil spils.

Dr Gerald T. Boalch,
11 Torridge Close,
Plymouth, PL7 2DH.

Peter H. van Bragt

Peter H. van Bragt is a molecular biologist with marine biology as a hobby. He has a great interest in the marine flora and fauna of the Netherlands with a special interest in Dutch sea slugs. He owns a large collection of photographs of Dutch marine species and habitats. He is strongly involved in Dutch marine biology monitoring projects and courses.

Peter H. van Bragt,
Antiloopstraat 28,
4817 LC BREDA,
The Netherlands.

Mark Breckels

Mark is a marine ecologist with an interest in trophic interactions and climate change. He is a keen photographer.


John Brooks

John Brooks is a keen angler and works for Angling-school.

John Brooks
Tel: 01209 314164

Greg Brown

Greg Brown was actively engaged in marine biological research, particularly nudibranch taxonomy, between 1970 and 1985, contributing to various publications and books intended to clarify identification of N.Atlantic species. He now manages oceanographic surveys conducted by Gardline Environmental Ltd.

Greg Brown
Forge Cottage
125 Yarmouth Road
Norwich NR7 0QY

Simon Brown

Simon Brown takes a wide range of photographs from shallow reefs to deep wrecks in temperate and tropical waters. He would be pleased to hear about any interesting projects or unusual requirement that users would like captured on film.

Simon Brown,
11 Cherberry Close,
Hants, GU51 2TD.

Francis Bunker

Francis Bunker has a large library of still and moving environmental images specialising in underwater photography around the British Isles. For more details visit:

Francis Bunker
Estuary Cottage
Pembrokeshire SA71 5RN
Wales, United Kingdom.
Tel (land line): +44 (0)1646 621277
Tel (mobile):   +44 (0)7968 486256
Check out the British Isles Seaweed Images Website

Ingvar Byrkjedal

Ingvar Byrkjedal
University of Bergen - Museum of Zoology
Museplass 3, N-5007 Bergen

Gerry Casey

Gerry Casey is the course organizer/co-ordinator for the Marine Conservation Society South East Group and is a keen amateur underwater photographer and member of Bsoup.

Gerry Casey,
11 Southfield Avenue,
Hertfordshire, WD24 7DT

Crown copyright, reproduced with the permission of CEFAS, Lowestoft

The Centre for Environment, Fisheries and Aquaculture Science (CEFAS) is a UK-based research and consultancy centre. CEFAS provides services in fisheries science and management, environmental monitoring and assessment, fish farming, fish health, disease and hygiene to clients around the world.

CEFAS Lowestoft Laboratory,
Pakefield Road,
Suffolk NR33 0HT

Stephen Chalk

Stephen is a catch and release angler, and is also studying marine biology.

Stephen Chalk

Nigel Charles

Nigel Charles holds images of saline lagoon and mollusc species but does not normally offer picture agency arrangements. MarLIN should be contacted if you wish to use commercially any of the images provided by Nigel Charles and included on the MarLIN Website.

Geoff Coates


Ken Collins

Dr Ken Collins is a marine scientist working at the Southampton Oceanography Centre with interests in artificial reefs and south coast marine habitats and species.


Dr Elizabeth Cook

Dr Elizabeth Cook is a marine biologist at the Scottish Association for Marine Science. Her main research interests are invasive species, new species for aquaculture and integrated aquaculture.

Scottish Association for Marine Science (SAMS)
Dunstaffnage Marine Laboratory
PA37 1QA
Tel: 01631 559 243
Fax: 01631 559 001

Countryside Council for Wales

The intertidal survey team of the Countryside Council for Wales has a collection of images marine species and habitats and site views and landscapes of the intertidal zone for the whole of Wales.

Mr Paul Brazier
Intertidal ecologist
Countryside Council for Wales,
Maes y Ffynnon,
LL57 2DW
Tel: 01248 387165

Inês Cristovão

Inês Cristovão is a marine biologist and Ph.D student at the University of Lisbon, specializing in the morphology and geometric morphometrics of decapods and gastropods, and population genetic differentiation.

Laboratório Marítimo da Guia (Faculdade de Ciências da Universidade de Lisboa),
Estrado do Guincho,
2750-374 Cascais.

Fiona Crouch

Fiona Crouch joined the MarLIN team in April 2006 to run the Shore Thing project. She has previously worked as the ranger for the St. Abbs and Eyemouth Voluntary Marine Reserve where her interest in underwater photography began.

MarLIN (as above)

Culture Collection of Algae and Protozoa (CCAP)

Image of specimen of Anotrichium barbatum (CCAP 1359/1) collected from an oyster bank near Pwllheli, Cymru by Dr C. Maggs in 1998.

The CCAP marine section is based at the Dunstaffnage Marine Laboratory, Scotland and the freshwater and protozoa section is based at CEH, Windermere. The CCAP is one of 10 microbial collections making up the National Culture Collection organisation ( The CCAP provides microbial and algal cultures for aquaculture, biotechnology, research and education.

Culture Collection of Algae and Protozoa
SAMS Research Services Ltd
Dunstaffnage Marine Laboratory
Argyll PA37 1QA
Tel: +44 (0)1631 559000
Fax: +44 (0)1631 559001

Mike Daines

Mike Daines is a professional photographer and photo-journalist, as well as being a keen diver.


As for Colin Speedie

Sue Daly

Sue Daly is a writer, broadcaster, underwater photographer and natural history film-maker based in the Channel Island of Jersey. Sue Daly is author of the book 'The Marine Life of the Channel Isles' (1998) and the 'Wildlife of the Channel Isles' Website. She holds a large collection of underwater photographs from the Channel Isles, the Red Sea, Maldives, Malaysia and the Mediterranean.

Sue Daly
La Vieille Maison,
La Seigneurie,
Sark GY9 0SF, Channel Islands

John Davenport

Professor John Davenport is a marine biologist who has worked on sea turtles for several years and has published papers on their swimming mechanisms and ecophysiology. He is particularly interested in turtle thermal regulation in relation to global warming and sex determination.

Department of Zoology and Animal Ecology
University College Cork
Lee Maltings
Prospect Row
Tel. (021) 904355

Misjel Decleer


Devon Wildlife Trust

Devon Wildlife Trust is the only independent organization exclusively concerned with all aspects of wildlife conservation in Devon.

Stephen Hussey
Devon Wildlife Trust,
Shirehampton House,
35-37 St David's Hill,
Exeter, EX4 4DA.

Kerry Donovan

Kerry Donovan gained a Marine and Freshwater Biology degree from the University of Wales, Aberystwyth in 2004. She is now a Primary School teacher.

23 Camden Street
PO12 3HZ

Dave Dunlop

Dave works for the Lancashire Wildlife Trust


Robert Durrant

Keen amateur, particularly interested in discovering the richness of shore life on the North Devon coast, having retired to that area of my birth

6 Clovelly Gardens South,
Devon, EX39 3DH

Richard Edmonds

Richard Edmonds holds a large collection of marine images from western Lyme Bay and a smaller Scottish and tropical collection. He also specialises in geology and fossils of the Dorset coast and gives illustrated, informal, non-technical lectures on all of these subjects.


Marco Faasse

Marco Faasse studies the distribution of marine invertebrates in The Netherlands. He is especially interested in certain underrecorded groups, e.g. Amphipoda, Ostracoda, Bryozoa, Hydrozoa, and in alien species. He owns a collection of images of species from these and other groups.

Marco Faasse
Schorerstraat 14
4341 GN Arnemuiden
The Netherlands

Liam Faisey

Marine life photographer and Angler currently studying a B.Sc. in Marine Biology at Plymouth University.

29 Upton Towans,
TR27 5BJ

David Fenwick Snr.

My interests lie in website design and wildlife photography and horticulture. I have a special interest in the photography of the Flora and Fauna of the Devon and Cornwall Peninsula. I am a very keen rockpooler, contributing images to Dr. Chris Gibson's recent book entitled Seashore.

84 Roscadghill Parc,
Heamoor, Penzance, Cornwall. TR18 3QY
Tel.: 01736 448392

Paula Ferris

Paula is Chair of Coastwise North Devon, and an amateur marine naturalist.

North Devon
EX33 1BP

Mike Firth

Mike Firth is a mature student and diver with an eye for photography. He has a limited supply of coastal photographs available.

16 Bransdale Gardens,
Leeds LS20 8QE

Alex Fraser

Alex Fraser is a technician in marine biology at the University of Plymouth and ready for any photographic opportunities.

Alex Fraser,
Room 703 Davy Building,
University of Plymouth,
Drake Circus,
Plymouth, PL4 8AA.

Guy Freeman

Guy Freeman is currently studying at Exeter University and has a keen interest in British marine life.



Food and Agriculture

Organization of the United Nations (FAO)

Images reproduced with permission from:

Roper, C.F.E., Sweeney, M.J. & Nauen, C.E., 1984. FAO species catalogue. Vol. 3. Cephalopods of the world. An annotated and illustrated catalogue of species of interest to fisheries. Food and Agriculture Organization of the United Nations, Rome, FAO Fisheries Synopsis, 125, 3, 277 pp.

Library enquiries
Viale delle Terme di Caracalla
00100 Rome, Italy
Tel: +39-6-57051
Fax: + 39-6-57053360

Terry Griffiths

Is a Torquay based diver and keen underwater photographer diving the South West dive sites.

89 Leeward lane

Dr Jason Hall-Spencer

Jason Hall-Spencer is a benthic ecologist specialising in calcareous marine organisms and underwater videography.

Jason Hall-Spencer
Royal Society Research Fellow
Marine Institute
University of Plymouth
Plymouth, PL4 8AA

Bryan Harrison

Bryan Harrison is a BSAC Sports Diver and photographer. He has been a member of Worthing BSAC since 2005.

Bryan Harrison
357 Upper Shoreham Road
West Sussex

Christian Henlé

Christian Henlé is a member of the Audio Visual Committee of the French Diving and Sub aquatic Activities Federation in Nice (FFESSM) and is a keen diver with a large collection of images of marine species and habitats mainly from around southern France.

Christian Henlé

John Hepburn

John Hepburn promotes the wider maritime sector - commercial, leisure and scientific - and is a member of the Finding Sanctuary local group. He also sails, dives and snorkels.

John Hepburn,
Mewstone Enterprises

Jacqueline Hill

Image redrawn by Jacqueline Hill from:

Sars, G.O.,  1890. Crustacea of Norway.  Volume 1, Amphipoda.  Christiania: ALB. Cammermayers Forlag.

Dr Hans Hillewaert

Hans Hillewaert,
Institute for Agriculture and Fisheries Research,
Flemish Government
Tel. +32-(0)59-56 98 32
Fax. +32-(0)59-33 06 29

Dr Hilmar Hinz

Hilmar Hinz graduated in Marine Biology from Newcastle in 1994, undertook a Masters Degree in Biology at the University of Oldenburg (Germany) and worked for the Senckenberg Institute (Wilhelmshaven, Germany). In 2001 he joined the Coastal Resource Ecology and Management group lead by Dr. M.J. Kaiser in Bangor (Wales) at the School of Ocean Sciences. In 2005 he completed his part-time PhD on the habitat use of flatfishes. He is currently working at the School of Ocean Sciences as a fisheries scientist.

Dr. Hilmar Hinz
School of Ocean Sciences
University of Wales, Bangor
Menai Bridge
LL59 5AB, UK
Tel + 44 (0)1248 388124

Dr Keith Hiscock

Dr Keith Hiscock has a large collection of images of marine species and habitats, landscapes, activities and impacts mainly from around Britain.

Keith Hiscock

Dr Sebastian P. Holmes

Dr Sebastian Holmes is a marine biologist who currently works at the Port Erin Marine Laboratory. His research interests include population genetics, intertidal ecology, invertebrate ecophysiology and supply side ecology.

Dr Sebastian Holmes

Davy Holt

Davy Holt is a recreational angler actively involved with the Glasgow Kelvingrove Museum's Common Skate Tagging Programme and also with the Common Skate Biodiversity Action Plan.

Davy Holt (preferred contact address).

Dr Rohan Holt

Rohan Holt works for the Countryside Council for Wales (CCW) and in his spare time Marine Wildlife Surveys (MWS). MWS is a small company specialising in underwater and marine biological photography, videography, species identification and Seasearch courses and benthic surveys.

Rohan holt,
Tan Hwfa,
North Wales, LL57 3LA.

Charles Hood

Charles Hood is a professional underwater photographer and journalist. He currently is contracted to DIVE magazine in the UK. He has over 12,000 UW images available for publication.

Charles Hood
Tel. 0771 262 2440

Tammy Horton

Dr Tammy Horton is undertaking a BP-funded fellowship on deep-sea amphipods at the Southampton Oceanographic Centre. She does not normally offer picture agency agreements.

Dr Tammy Horton,
BP Fellow UK,
National Oceanography Centre, Southampton
George Deacon Division
Waterfront Campus, European Way
Southampton SO143ZH
Tel: (02380) 596352
Fax: (02380) 596247

Mike Hosken

Mike Hosken is a 'retired agriculturist, one-time sailor, quasi-oceanographer and seaside dog-walker'.

Mike Hosken,

Martin Isaeus

Martin Isaeus is a plant ecologist working in the Deptartment of Botany, Stockholm University (

Martin Isaeus
Norwegian Institute of Water Research NIVA,
Brekkeveien 19,
Postboks 173 Kjelsaas,
0411 Oslo,
Tel: (+47) 22 18 51 77
Fax: (+47) 22 18 52 00

Angus Jackson

Dr Angus Jackson is a Research Fellow in marine ecology at the Environmental Research Institute of North Highland College at the University of the Highlands and Islands. (

Environmental Research Institute,
North Highland College, UHI
Ormlie Road,
Caithness, KW12 7EE.

Norman Jackson

Norman Jackson is a yachtsman and amateur naturalist. He has a collection of images, primarily of the marine life you see out at sea from the surface around the north west of Scotland.

Please email the MarLIN ( team directly if you wish to use any of Norman Jackson images we have online.

Matthew Jeanes

Matt is a freediver and spearfisherman who shoots very few fish and spends a lot of time taking photos. He is building a comprehensive collection of British marine life photos.


Joint Nature Conservation Committee

The Joint Nature Conservation Committee (JNCC) is the UK Governments national and international wildlife advisor. JNCC holds a large collection of seabed marine species, habitat and location images mainly from surveys undertaken by the Marine Nature Conservation Review. JNCC is not a picture agency but should be contacted if you wish to use commercially any of the images provided by JNCC and included on the MarLIN Website.

Communications Team,
Joint Nature Conservation Committee,
Monkstone House,
Peterborough, PE1 1JY, UK
Tel: +44 (0)1733 562626
Fax: +44 (0)1733 555948

Dick Jones

Image reproduced with the authors permission from:

Legg, G. & Jones, R. E., 1988. Pseudoscorpions. Synopses of the British Fauna (New Series) (ed. D.M. Kermack & R.S.K. Barnes), The Linnean Society of London. Avon: The Bath Press. [Synopses of the British Fauna No. 40.]

Dick Jones has a large collection of images of creepy crawlies and other natural history objects

RE Jones,
14, Post Office Road,
King's Lynne,
Norfolk, PE31 6HP

Dylan Jones


Rob Jutsum

Rob Jutsum is an amateur photographer. He has several hundred images of rocky shore animals which he uses to support the activities of "Coastwise", North Devon.

36 Stallards,
Devon EX33 1BP

Jenni Kakkonen

Jenni Kakkonen is a biologist for "Marine Services" at Orkney Islands Council and as part of her job conducts intertidal rocky shore surveys. She is also a keen diver.

Jenni Kakkonen
Marine Services
Harbour Authority Building
Scapa, Orkney
KW15 1SD

Robert Keen

Robert J. Keen is a keen diver who enjoys observing, photographing and cataloging as much marine life as he can from around the world. He is passionate about stopping the degradation of the Oceans particularly as regards the Shark populations.

Robert J. Keen
2 Dellfield Close,
Herts. WD17 4LB

John Kelly

John Kelly is a Research Associate for Quercus, Queens University of Belfast.

John Kelly,
Quercus Research Associate,
Queen's University of Belfast,
Medical Biology Centre,
97 Lisburn Road,
Belfast BT9 7BL
Northern Ireland .
Tel: +44 (0) 2890 972 259

Hans Kleef and Peter Tydeman

Hans Kleef and Peter Tydeman are marine ecological technicians at the National Institute for Coastal and Marine Management/RIKZ, The Netherlands.

National Institute for Coastal and Marine Management / RIKZ 207 9750
AE Haren
The Netherlands
T: (0)31 50 533 13 78 and

Dr Stefan Kraan

Dr Stefan Kraan is a research scientist working at the Martin Ryan Marine Science Institute who holds images of some seaweed species but does not normally offer picture agency agreements.

Irish Seaweed Industry Organisation
Martin Ryan Marine Science Institute
National University of Ireland, Galway
Galway Ireland
Fax: 353-91-525005

Dr Inken Kruse

Dr Inken Kruse is a marine research scientist, mainly on population genetics issues of marine invertebrates, who carried out her PhD on Scoloplos armiger.

Dr Inken Kruse,

Holly Latham

Holly Latham is a marine biology graduate, MarLIN and MBA volunteer and SCUBA instructor, with personal interests covering a variety of anthropogenic marine impacts. She particularly loves macro photography and aims to take photographs that capture the texture and vibrancy of our coastline.


Gordon Lang

Gordon is a web and graphic designer, photographer and diver.


Jack Laws

Jack Laws has been diving for over 50 years and has underwater and landscape images of marine reserves in Scotland, Bonaire and Sulawesi.

Jack Laws
High street,
Eyemouth, TD14 5NL

Dan Lear

Dan Lear is the Information Systems Co-ordinator with the MarLIN team.

MarLIN (as below)

Gerald Legg

Gerald Legg is the natural historian at the Booth Museum of Natural History, Brighton, natural history photographer and author of childrens' books.

20 Manor Gardens,
West Sussex, BN6 9UG
Pseudoscorpion in Britain

Paula Lightfoot

Paula Lightfoot is a tutor for the Marine Conservation Societyís Seasearch recording scheme and records officer for the Yorkshire Naturalistsí Unionís marine and coastal section.

Paula Lightfoot,
Seasearch North East,

For information on Seasearch courses and dives visit:
For information on the Yorkshire Naturalistsí Union marine and coastal section visit:

Richard Lord

Richard Lord is a professional marine photographer with a large collection of images of marine life from around Guernsey and commercial fish and shellfish from around the world.

Richard Lord
Guernsey, GY1 1BQ,
Great Britain.
Fax: +44 (0)1481 700699

Dr Sutherland MacIver

Dr Sutherland MacIver is a senior lecturer of biochemistry at the University of Edinburgh and keen scuba diver.


Julian Madle

An avid scuba diver with a lifelong interest in wildlife and nature photography, both terrestrial and undersea.

1-2 New Road,
Middleton by Wirksworth,
Derbyshire, DE4 4NA


Dr Christine Maggs

Dr Christine Maggs is a phycologist studying all aspects of marine red and green algae, and specializing in taxonomy of red algae. She is co-author of the Ceramiales (red algae) flora of the British Isles, published by the Natural History Museum.

School of Biology & Biochemistry
Queen's University of Belfast
Belfast BT9 7BL
Northern Ireland.
Tel: 02890 972265

Marine Biological Association of the United Kingdom (MBA)

The Marine Biological Association of the United Kingdom (MBA) was founded in 1884 and its laboratory on Citadel Hill, Plymouth opened in 1888. The aims of the Association are to promote scientific research into all aspects of life in the sea and to inform the public of the knowledge gained. The MBA is an independent research association supported by grant in aid from NERC and a Registered Charity. The MBA has a small collection of images belonging specifically to the Association including historical material. The MBA is not a picture agency but should be contacted if you wish to use commercially any of the images provided by the MBA and included on the MarLIN Website.

Directors Secretary
Marine Biological Association of the United Kingdom,
The Laboratory,
Citadel Hill,
Plymouth, PL1 2PB, UK.
Tel: +44 (0)1752 633336
Fax: +44 (0)1752 633102

Marine Biodiversity and Climate Change - MarClim

MarClim is a new major four-year multi-partner British and Irish project led by the Marine Biological Association in collaboration with Plymouth Marine Laboratory, The Scottish Association for Marine Science, The University of Plymouth and University College Cork. The aim of MarClim is to assess and forecast the influence of climatic change on marine biodiversity in Britain and Ireland using long term and current data on intertidal rocky shore indicator species. MarClim is currently the only government-funded project addressing climate change in the marine environment in the UK.

The MarClim Programme
The Marine Biological Association of the UK
The Laboratory, Citadel Hill
Plymouth, PL1 2PB
Devon, UK
Tel: +44(0)1752 633287
Fax: +44(0)1752 633102

Marine Conservation Society

Calum Duncan has a collection of MCS copyright digital and scanned images depicting subtidal marine life, coastal scenes and activities that he has taken throughout Scotland. Please contact him at the MCS Scotland Offices for further information.

Calum Duncan
Scottish Conservation Manager
Marine Conservation Soceity
3 Coaates Place,
Tel: +44 (0) 131 226 6360
Fax: +44 (0) 131 226 2391

Marine Life Information Network for Britain and Ireland (MarLIN)

MarLIN is an initiative of the Marine Biological Association of the UK in collaboration with major holders and users of marine biological data. MarLIN will provide the most comprehensive and easily used source of information about marine habitats, communities and species around Britain and Ireland.

Marine Biological Association of the United Kingdom,
The Laboratory,
Citadel Hill,
Plymouth, PL1 2PB,
Tel: +44 (0)1752 633336
Fax: +44 (0)1752 633102

Mike Markey

Mike is a Poole-based diver and dive charter boat skipper with an interest in underwater photography.

Mike Markey,
20 Warwick Road,
Southampton, SO40 3QP.
Tel: 07732 271017

Andrew Marriott

Andrew Marriott is a marine biologist, keen marine conservationist and wildlife photographer.

Andrew Marriott
14 Drayton Street,
West Midlands,WS2 9QB

Charlotte Marshall

Charlotte Marshall is a member of the MarLIN team and an amateur photographer with an interest in the conservation of marine ecosystems and fisheries.

MarLIN (as above)

Derek Mayes

Derek Mayes lives overlooking Scapa Flow (Orkney). Mainly a photographer but with an interest in all aspects of Natural History.


Cathal McNaughton


Ruth Memmott

Ruth is a marine biologist studying for a masters at the University of Plymouth.


Christopher McAskie

Christopher McAskie is a retired and amateur photographer of shorelife and wildlife.

24 Penydre Neath
SA11 3HD

Jon Moore

Jon Moore, freelance marine environmental consultant, has a large collection of slides and a rapidly increasing collection of digital photos of subtidal, intertidal and coastal species, habitats, resources and impacts from UK and overseas.

Jon Moore,
T? Cara, Point Lane,
Cosheston, Pembroke Dock,
Pembrokeshire, SA72 4UN,
Wales, UK.

Natural History Museum (NHM)

The Natural History Museum (NHM) promotes the discovery, understanding, enjoyment, and responsible use of the natural world.

Paul Clark,
Natural History Museum,
Cromwell Road,
London SW7 5BD, UK.
Tel: +44 (0)20 7942 5000

National Museum Wales (NMW)

National Museum Wales (NMW) holds internationally renowned collections covering the art, archaeology, botany, geology, industry, social history and zoology of Wales. Through its collections, they explore Wales and its place in the world.

Anyone wishing to use their images for any reason must apply to the museum in writing for permision.

Kay Kays
Image Licensing Officer,
National Museum Wales,
Cathays Park,
Cardiff, CF10 3NP.
Tel: (029) 2039 7951

Paul Naylor

Paul Naylor is a marine biologist, underwater photographer and author of the book 'Great British Marine Animals'. He has a large collection of marine animal images from around the British Isles.

Tel: 07041 - 351307

Paul Newland

Paul Newland is a widely traveled diver for many years and currently a member of the Totnes Subaqua Club with a keen interest in marine life.

Paul Newland
Tel: 01364 642287

Tim Nicholson

Tim Nicholson is an amateur diver and underwater photographer. He has a collection of images taken in several locations around the world all on his trusty Nikonos V. Most of his British images were taken around the Isle of Man, where he currently lives and works. He is a contributor to photo galleries at

42 Garth Avenue,
Isle of Man, IM9 6QZ
Tel: 01624 836067

Thom Nickell

Thom Nickell is a benthic ecologist based at the Scottish Association for Marine Science.

Thom Nickell
Scottish Association for Marine Science
Argyll, PA37 1QA
Tel: 01631 559261

Helmut Nisters

Helmut Nisters is curator of the shell collection of the Natural History Department of the Tiroler Landesmuseum Ferdinandeum, Innsbruck.

Helmut Nisters,
Franz-Fischer-Str. 46,
A-6020 Innsbruck,

Peter O'Donnell

Peter O'Donnell works for the Loughs Agency and is involved in its environmental education programme. He has a collection of images of species native to the UK.

Peter O'Donnell,
22 Victoria Road,
BT47 2AB
Tel: (028) 71342100

Judith Oakley

Judith Oakley, MSc, BSc specialises in macro intertidal rocky shore photography. She contributes many images to marine conservation charities and has had work published by the Countryside Council for Wales, Defra, Marine Biological Association and BBC Wildlife magazine. She has a keen interest in seahorses and pipefish, marine intertidal species, sea slugs, marine mammals and marine conservation issues in general.


Paul Parsons

Paul Parsons is a marine photographer with a large collection of images. Visit to view.

Paul Parsons,

Dave Peake

Dave Peake is an underwater photographer who has been diving for over 30 years. He specialises in photography in British waters.


Tabitha Pearman

Tabitha works in offshore marine environmental survey and enjoys rock pooling and snorkelling.


Andrew Pearson

Andrew Pearson has a background in marine ecology and has a large collection of coastal and underwater images of marine life and habitats.

Andrew Pearson,
46 Grove Road,
SO15 3GG, UK.
Tel: 07905 607 027

Paul Peek

Paul Peek takes photos on behalf of the Northfield St. Nicholas Primary School.

Paul Peek,
Northfield St. Nicholas Primary School,
St Margaret's road,
Suffolk NR32 4HN,

Bernard Picton

Bernard Picton has a large collection of colour slides of marine species and habitats from around Britain, Ireland, the Mediterranean, Caribbean and Indo-Pacific with a special interest in nudibranch molluscs and sessile invertebrates.


Anne Marie Power

Anne Marie Power has collected digital images of intertidal and shallow subtidal organisms on Irish coasts while conducting surveys for the MarClim project.

Anne Marie Power
Department of Zoology,
Room 226,
Martin Ryan Institute,
National University of Ireland,
Galway, Ireland
Tel: 353 91 493015
Fax: 353 91 525005

Dr Ian Powell

Dr Ian Powell is a research scientist working at Edge Hill College of Higher Education. He has a large collection of images relating to ecology, conservation and landscapes around the world.

Dr Ian Powell,
Natural and Applied Sciences,
Edge Hill,
St Helen's Road, Ormskirk,
Lancashire, L39 4QP.

John Rafferty

John is a leading marine and landscape photographer and is also a photo/journalist for the Irish and UK fishing industry. A selection of his images can be viewed on his website.

John Rafferty,
Co. Donegal, Ireland.

Fiona Ravenscroft

Seasearch Surveyor E-mail:

Ivor Rees

Ivor Rees has a collection of seabed habitat photos from various parts of the Irish Sea taken with cameras mounted on seabed sledges.

Ivor Rees,
c/o School of Ocean Sciences,
University of Wales, Bangor
Menai Bridge
Anglesey, Gwynedd LL59 5EU
Tel: 01248 382846

Abigail Rhodes

Abigail Rhodes is an artist and wildlife enthusiast.

Abigail Rhodes,
78 Dalcharn,
Nr Lairg,
Sutherland IV27 4XU

Graham Rickett

I have 33 years experience in research and development in antibacterials, antivirals, antifungals and an experienced GPCR biologist as well as competent in new technology research. As a volunteer for the Thanet Coastal Project, using my marine biology background, I am hoping to map West Bay and Epple Bay, Kent for all fauna and flora throughout the littoral zone and cliff areas of the location eventually.

Graham Anthony Rickett,
9 Ursuline Drive,
Kent CT8 8HX,

Dr J. Murray Roberts

Dr J. Murray Roberts is a Reader in Biodiversity at Heriot-Watt University working on the biology and ecology of cold-water corals.

School of Life Sciences,
Herriot-Watt University,

Dan Ros

Dan is PADI Master Scuba Diver and enjoys macro photography and fish portraits.


John Rundle

John Rundle has a collection of images of marine species, including large numbers of cuttlefish images.

John Rundle,
Marine Biological Association of the UK,
Tel: 01752 633350

Harriet Salomonsen

Harriet is a marine ecologist currently based in Southampton. She is also a keen photographer.

Harriet Salomonsen,
Church close,
Nr Tiverton,
Devon, UK.

Nigel Sawyer

Nigel Sawyer,
14 High Ashton,
119 Kingston Hill,
Kingston upon Thames,
Surrey KT2 7QL, UK.

Scottish Association for Marine Science (SAMS)

Dunstaffnage Marine Laboratory (DML) near Oban, Argyll is the home of the Scottish Association for Marine Science (SAMS). SAMS carries out research in coastal and offshore environments, with a particular emphasis on biology, sediment geochemistry and technology. Increasingly, results are being applied to environmental management/impact assessment. The University of the Highlands and Islands Degree Course in Marine Science is delivered from Dunstaffnage - see

Dunstaffnage Marine Laboratory,
Argyll, PA37 1QA,
Tel. 01631 559000
Fax. 01631 559001

Scottish Natural Heritage (SNH)

Scottish Natural Heritage (SNH) is a government agency that works to conserve and enhance Scotland's natural heritage of wildlife, habitats and lanscapes. SNH aims to help people enjoy Scotlands natural heritage responsibly, understand it more fully and use it wisely so that it can be sustained for future generations.

Scottish Natural Heritage (SNH),
Maritime Group,
2/5 Anderson Place,
Edinburgh, EH6 5NP.
Tel: 0131 446 2400
Fax: 0131 446 2405

Sue Scott

Sue Scott is a professional marine life photographer and biologist with a large collection of images especially from Scottish marine and freshwater habitats.

Sue Scott
Strome House
North Strome
Loch Carron
Ross-shire, IV54 8YJ, UK
Fax: +44 (0)1520 722588 has a large collection of images of UK marine fauna including an extensive collection of photo micrographs as well as images of marine surveys, laboratory work and activities associated with marine developments. They also hold a large amount of quantitative and qualitative data collected on surveys carried out all around the UK and spanning the last 30years.

Bryony Pearce
Contracts Director,
Marine Ecological Surveys Limited,
24a Monmouth Place,
Bath, BA1 2AY

Dennis R. Seaward

Dennis R. Seaward holds images of saline lagoon and mollusc species but does not normally offer picture agency arrangements. MarLIN should be contacted if you wish to use commercially any of the images provided by Dennis Seaward and included on the MarLIN Website.

Becky Seeley

Becky works as the Biological records officer for the Data Archive for Seabed Species and Habitats (DASSH) project at the MBA.


Guido Selling

Guido Selling is an underwater photographer with an interest in marine biology.

Guido M.A. Selling,
De Nesse 13,
2291 XN Wateringen,
The Netherlands.


The "Scientific and Environmental ROV Partnership using Existing iNdustrial Technology" (SERPENT) project aims to make cutting-edge ROV technology and data more accessible to the world's science community, sharing knowledge and progressing deep-sea research. The programme will interact with science and conservation groups globally and transparently communicate our project to the public to increase the awareness of our fragile marine resources.

SERPENT is a global project hosted by the DEEPSEAS group, in the George Deacon Division (GDD) of the National Oceanography Centre, Southampton (NOCS) with a network of UK and global partners.

Maria Baker,
ChEss Coordinator/DC-UK Research Fellow
DEEPSEAS Group, George Deacon Division,
National Oceanography Centre, Southampton, Empress Dock,
Southampton SO14 3ZH, UK.
Tel: +44 (0)23 80596352
Fax: + 44 (0)23 80596247

Dr Matthew Service

Dr Matthew Service is a Researcher employed by the Department of Agriculture and Rural Development for Northern Ireland. He has conducted a number of seabed surveys in the Irish Sea and Coastal waters of Northern Ireland for both fisheries and environmental studies.

Dr Matthew Service,
Agriculture and Environmental Science Division,
Department of Agriculture and Rural Development,
Newforge Lane,
Tel: 44 (0)28 90255502
Fax: 44 (0)28 90255004

Jack Sewell

Jack Sewell is a member of the MarLIN team, an illustrator and amateur photographer with an interest in environmental education, marine species and habitats.

MarLIN (as above)

Tim Siferd

Tim Siferd is a biologist in the Stock Assessment section of Fisheries and Oceans Canada. Artic benthic stocks, especially clams are assessed through remote underwater photographic sampling and diving.

Tim Siferd,
Fisheries and Oceans Canada,
501 University Crescent,
Winnipeg, MB, Canada,
R3T 2N6.

Professor Alan Southward B.Sc., Ph.D., D.Sc.

Professor Southward (now deceased) was an honoury research fellow at the Marine Biological Association of the United Kingdom. His research interests included long-term changes in marine ecosystems; the biogeography and systematics of Cirripedia, and the biology of hydrothermal vents and seeps.

For image enquiries contact:

Marine Biological Association of the United Kingdom,
The Laboratory,
Citadel Hill,
Plymouth, PL1 2PB, UK.
Tel: +44 (0)1752 633336
Fax: +44 (0)1752 633102

Ian F. Smith 

Ian Smith is interested in seashore invertebrates and macrophotography of living specimens in water. He contributes images and articles to the Conchological Society of G.B. & Ireland’s magazine, ‘Mollusc World’, and online Encyclopedia of Molluscs.

12 Fernhill, Mellor, Stockport, SK6 5AN


Dr Richard Smith

Richard Smith is an entomologist with special interests in invertebrate conservation and natural history education.

Dr Richard Smith,
12 Chestnut grove,

Colin Speedie

Colin Speedie is a professional yachtsman with many years experience conducting marine life surveys afloat, and is currently running a basking shark research project on behalf of the Joint Marine Programme of The Wildlife Trusts and WWF-UK. He has a wide variety of surface images of marine species from around the UK. Colin Speedie is not a picture agency but should be contacted if you wish to use commercially any of the images provided by him and included on the MarLIN Website.

Colin Speedie,
Seawatch Charter,
Waterside House,
Falmouth Road,
Penryn, Cornwall, TR10 8BE
Tel/Fax: +44 (0) 1326 373360

Cathy Stevenson

Cathy Stevenson is an amateur rockpooler and photographer. The photos were taken as part of the Sealife Survey (


Dr Matthias Strasser

Dr Matthias Strasser is a marine research scientist working at the Wadden Sea Station Sylt of the Alfred Wegener Institute Foundation for Polar and Marine Reasearch (AWI). However, he does not normally offer picture agency arrangements.

Dr Matthias Strasser,
Wadden Sea Station Sylt,
Alfred Wegener Institute Foundation for Polar and Marine Research,
Hafenstrasse 43,
25992 List/Sylt.

Erling Svensen

Erling Svensen has been an underwater photographer for 28 years with a speciality on marine life and more than 40,000 photographs in stock.

Ocean Photo - Erling Svensen,
Hamraneveien 16
4370 Egersund - Norway

Mark Thomas

Mark Thomas is an amateur diver with the Hartford SAC, diving since 1998 and taking underwater photographs since 2002 (using a Nikon F90X camera). Mark dives mostly around Ireland, Scotland and Wales.

Mark Thomas,
38 The Crescent,
Cheshire, CW9 8AD.

Alan Thurbon

As the result of a guided walk by Bill Farnham of Portsmouth University, Alan Thurbon is discovering the surprising diversity of sea life underneath Langstone Bridge, which connects to Hayling Island.


Pamela Tompsett

Pamela Tompsett is Coordinator of the Helford River Voluntary Marine Conservation Area and a keen biologist. However, she does not normally enter into picture agency agreements. MarLIN should be contacted if you wish to use commercially any of the images provided by Pamela Tompsett and included on the MarLIN Website.

Paul Tranter

Paul Tranter works for the Sir Alister Hardy Foundation for Ocean Science as a plankton analyst. From 1958-88 Paul was employed by the Marine Biological Association of the United Kingdom and during that time he developed a keen interest in corals and sea anemones and made many observations on behavioural activity.

Paul Tranter
Sir Alister Hardy Foundation for Ocean Science (SAHFOS),
The Laboratory,
Citadel Hill,
PL1 2PB.
Tel +44 (0) 1752 633288

Jo Trout

Jo Trout,
Silver Dolphin Centre,
The Old Lifeboat House,
Mount Pleasant Road,
Cornwall, TR13 9JS.

Harvey Tyler-Walters

Harvey Tyler-Walters is the Senior Data Researcher with the MarLIN team and an amateur photographer with an interest in marine intertidal species and coastal actvities.

Email: Harvey Tyler-Walters

Sofie Vandendriessche

Sofie Vandendriessche is a PhD student studying the fauna associated with detached seaweeds floating at the sea surface in Belgian coastal waters.

Sofie Vandendriessche, MSc
Ghent University, Marine Biology Section

Dr Leen Vandepitte

Research Interests:

Dr Leen Vandepitte,
Vlaams Instituut voor de Zee,
(Flanders Marine Institute),
Tel. +32-(0)59-34 01 55
Fax. +32-(0)59-34 21 31

Bryan Wasson

Bryan Wasson is a freelance consultant specialising in the taxonomic identification of marine and estuarine benthic macrofauna.

Bryan Wasson,
6 King St,
PL22 0AQ
Tel: +44 (0) 1208 871283

Peter Watkinson

Peter Watkinson is a commercial fisherman and can supply images and specimens of many North Sea species.

Peter Watkinson

Rob Spray and Dawn Watson

Rob Spray and Dawn Watson have a large collection of images of marine species and habitats from Britain and around the world, many taken while working as joint coordinator for Seasearch in East Anglia.

Rob Spray and Dawn Watson
1 Town Houses,
Yoxford Road
Suffolk IP17 2LX

Whirling Disease Foundation

The Whirling Disease Foundation, Inc. is a non-profit corporation established in 1995 with a mission to raise the funds needed for researched solutions to the damage caused by whirling disease. The Whirling Disease Foundation has images of whirling disease-infected fish, Myxobolus cerebralis (the parasitic cause of whirling disease), and Tubifex tubifex (the parasite's alternate host).

Whirling Disease Foundation
P.O. Box 327
Bozeman , Montana 59771-0327
(406) 585-0860

Kristie Wilkinson

Kristie Wilkinson is an underwater wildlife photographer.

Kristie Wilkinson

Bob Williams

Bob Williams is an Honorary Research Fellow at Plymouth Marine Laboratory.

Bob Williams,
Myrtos Porthrepta Road,
Carbis Bay,
St Ives,
Cornwall TR26 2LB

Kate Willis

Kate Willis is a Marine Ecologist at the Scottish Association for marine science. Research interests are zooplankton ecology and invasive species.

Scottish Association of Marine Science
Argyll, PA37 1QA
Tel: 01631 559313

Douglas P. Wilson

Images reproduced with the permission of the Journal of the Marine Biological Association of the UK from:

Wilson, D.P., 1929-30. The larvae of the British Sabellarians. Journal of the Marine Biological Association of the UK, 16, 221-270.

Executive editor,
Journal of the Marine Biological Association of the UK,
The Laboratory,
Citadel Hill,
Plymouth, PL1 2PB.

P. Terry Wimbleton

P. Terry Wimbleton,
5 Ramsdale Avenue,
Leigh Park,
Havant, Hants, PO9 4DY.

Ms Jessica Winder

Jessica Winder is a keen photographer with a collection of images of marine species from shores around the United Kingdom


Chris Wood

Chris Wood is the organizer of the Marine Conservation Society's (MCS) annual programme of dives and diving surveys and has a large number of images of British, Red Sea, Maldivian and Malaysian species and habitats, mostly taken on MCS surveys.


Tim Worsfold

Tim Worsfold is the Consultancy and Research Manager at Thomson Unicomarine; involved in marine ecology reporting, identification of specimens from benthic samples and in situ, as well as shore trips outside work.

Tim Worsfold,
Thomson Unicomarine Ltd
7 Diamond Centre, Works Road
Letchworth, Herts, SG6 1LW, UK
Tel. +44 (0)1462 675559
Fax. +44 (0)1462 483103 <> <>

WWF - The Environmental Network

WWF is a worldwide conservation network and its photo library reflects the scope of its work with 20,000 colour transparencies on a range on a range of UK and international conservation and environmental issues. Categories include mammals, birds, land/water invertebrates, UK and international habitats/scenery and a wide range of images depicting environmental issues. For more information about the use of these images, please contact the WWF photo-library.

Tel: +44 (0) 1483 426444 or
The library is open between 9am and 5pm.

Dr Russell B Wynn

Dr Russell Wynn is a Senior Marine Scientist at the National Oceanography Centre, Southampton. He is also an Associate Researcher at SAHFOS, and is co-ordinator of the SeaWatch SW project (

Dr Russell B Wynn
Senior Marine Scientist
National Oceanography Centre
European Way
Southampton, SO14 3ZH
Tel: +44 (0)23 80596553 (office)
80596555 (secretary)
80595000 (switchboard)